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Tuesday, June 15, 2010

Dolphin

For other uses, see Dolphin (disambiguation). This article is semi-protected. Bottlenose Dolphin breaching in the bow wave of a boat. Dolphins are marine mammals that are closely related to whales and porpoises. There are almost forty species of dolphin in seventeen genera. They vary in size from 1.2 m (4 ft) and 40 kg (90 lb) (Maui's Dolphin), up to 9.5 m (30 ft) and 10 tonnes (9.8 LT; 11 ST) (the Orca or Killer Whale). They are found worldwide, mostly in the shallower seas of the continental shelves, and are carnivores, mostly eating fish and squid. The family Delphinidae is the largest in the Cetacean order, and relatively recent: dolphins evolved about ten million years ago, during the Miocene. Dolphins are among the most intelligent animals and their often friendly appearance and seemingly playful attitude have made them popular in human cultu

Origin of the name

The name is originally from Ancient Greek (delphís; "dolphin"), which was related to the Greek (delphys; "womb"). The animal's name can therefore be interpreted as meaning "a 'fish' with a womb". The name was transmitted via the Latin delphinus, Middle Latin dolfinus and the Old French daulphin, which reintroduced the ph into the word. The word is used in a few different ways. It can mean: * Any member of the family Delphinidae (oceanic dolphins), * Any member of the families Delphinidae and Platanistoidea (oceanic and river dolphins), * Any member of the suborder Odontoceti (toothed whales; these include the above families and some others), * Used casually as a synonym for Bottlenose Dolphin, the most common and familiar species of dolphin. This article uses the second definition and does not describe Porpoises (suborder Odontoceti, family Phocoenidae). Orcas and some closely related species belong to the Delphinidae family and therefore qualify as dolphins, even though they are called whales in common language. A group of dolphins is called a "school" or a "pod". Male dolphins are called "bulls", females "cows" and young dolphins are called "calves".

Taxonomy

Suborder Odontoceti, toothed whales Family Delphinidae, oceanic dolphins Genus Delphinus Long-Beaked Common Dolphin, Delphinus capensis Short-Beaked Common Dolphin, Delphinus delphis Genus Tursiops Common Bottlenose Dolphin, Tursiops truncatus Indo-Pacific Bottlenose Dolphin, Tursiops aduncus Genus Lissodelphis Northern Rightwhale Dolphin, Lissodelphis borealis Southern Rightwhale Dolphin, Lissodelphis peronii Genus Sotalia Tucuxi, Sotalia fluviatilis Costero, Sotalia guianensis Genus Sousa Indo-Pacific Hump-backed Dolphin, Sousa chinensis Chinese White Dolphin (the Chinese variant), Sousa chinensis chinensis Atlantic Humpbacked Dolphin, Sousa teuszii Genus Stenella Atlantic Spotted Dolphin, Stenella frontalis Clymene Dolphin, Stenella clymene Pantropical Spotted Dolphin, Stenella attenuata Spinner Dolphin, Stenella longirostris Striped Dolphin, Stenella coeruleoalba Genus Steno Rough-Toothed Dolphin, Steno bredanensis Genus Cephalorhynchus Chilean Dolphin, Cephalorhynchus eutropia Commerson's Dolphin, Cephalorhynchus commersonii Heaviside's Dolphin, Cephalorhynchus heavisidii Hector's Dolphin, Cephalorhynchus hectori Genus Grampus Risso's Dolphin, Grampus griseus Genus Lagenodelphis Fraser's Dolphin, Lagenodelphis hosei Genus Lagenorhynchus Atlantic White-Sided Dolphin, Lagenorhynchus acutus Dusky Dolphin, Lagenorhyn chus obscurus Hourglass Dolphin, Lagenorhynchus cruciger Pacific White-Sided Dolphin, Lagenorhynchus obliquidens Peale's Dolphin, Lagenorhynchus australis White-Beaked Dolphin, Lagenorhynchus albirostris Genus Orcaella Australian Snubfin Dolphin, Orcaella heinsohni Irrawaddy Dolphin, Orcaella brevirostris Genus Peponocephala Melon-headed Wha le, Peponocephala electra Genus Orcinus Killer Whale (Orca), Orcinus orca Genus Feresa Pygmy Killer Whale, Feresa attenuata Genus Pseudorca False Killer Whale, Pseudorca crassidens Genus Globicephala Long-finned Pilot Whale, Globicephala melas Short-finned Pilot Whale, Globicephala macrorhynchus Genus †Australodelphis †Australodelphis mirus Family Platanistidae Ganges and Indus River Dolphin, Platanista gangetica with two subspecies Ganges River Dolphin (or Susu), Platanista gangetica gangetica Indus River Dolphin (or Bhulan), Platanista gangetica minor Family In iidae Amazon River Dolphin (or Boto), Inia geoffrensis Family Lipotidae Baiji (or Chinese River Dolphin), Lipotes vexillifer (possibly extinct, since December 2006) Family Pontoporiidae La Plata Dolphin (or Franciscana), Pontoporia blainvillei Six species in the family Delphinidae are commonly called "whales" but genetically are dolphins. They are sometimes called blackfish. Melon-headed Whale, Peponocephala electra Killer Whale (Orca), Orcinus orca Pygmy Killer Whale, Feresa attenuata Wolphin Kawili'Kai at the Sea Life Park in Hawaii. False Killer Whale, Pseudorca crassidens Long-finned Pilot Whale, Globicephala melas Short-finned Pilot Whale, Globicephala macrorhynchus

Hybrid dolphins

In 1933, three strange dolphins beached off the Irish coast; they appeare d to be hybrids between Risso's and Bottlenose Dolphins. This mating was later repeated in captivity producing a hybrid calf. In captivity, a Bottlenose Dolphin and a Rough-toothed Dolphin pro

duced hybrid offspring. A Common-Bottlenose hybrid lives at SeaWorld California. Other dolp hin hybrids live in captivity around the world or have been reported in the wild, s uch as a Bot tlenose- Atlantic Spotted hybrid. The best known hybrid is the Wolphin, a False Killer Whale-Bottlenose Dolphin hybrid. The Wolphin is a fertile hybrid. Two Wolphins currently live at the Sea Life Park in Hawaii; the first was born in 1985 from a male False Killer Whale and a female Bottlenose. Wolphins have also been observed in the wild.

Evolution and anatomy

Evolution


See also: Evolution of cetaceans Dolphins, along with whales and porpoises, are descendants of terrestrial mammals, most likely of the Artiodactyl order. The ancestors of the modern day dolphins entered the water roughly fifty million years ago, in the Eocene epoch. Hind Limb Buds on Dolphins. An embryo of a Spotted Dolphin in the fifth week of developme nt. The hind limbs are present as small b umps (hind limb buds) near the base of the tail. The pin is approximately 2.5 cm (1.0 in) long. Modern dolphin skeletons have two small, rod-shaped pelvic bones thought to be vestigial hind limbs. In October 2006 an unusual Bottlenose Dolphin was captured in Japan; it had small fins on each side of its genital slit which scientists believe to be a more pronounced development of these vestigial hind limbs.

Anatomy


Dolphins have a streamlined fusiform body, adapted for fast swimming. The tail fin, called the fluke, is used for propulsion, while the pectoral fins together with the entire tail section provide directional control. The dorsal fin, in those species that have one, provides

stability while swimming. Though it varies per species, basic coloration patterns are shades of grey usually with a lighter underside, often with lines and patches of different hue and contrast. The head contains the melon, a round organ used for echolocation. In many species, elongated jaws form a distinct beak; species such as the Bottlenose have a curved mouth which looks like a fixed smile. Some species have up to 250 teeth. Dolphins breathe through a blowhole on top of their head. The trachea is anterior to the brain. The dolphin brain is large and highly complex and is different in structure from that of most land mammals. Unlike most mammals, dolphins do not have hair, except for a few hairs around the tip of their rostrum which they lose shortly before or after birth. The only exception to this is the Boto river dolphin, which has persistent small hairs on the rostrum. Dolphin’s reproductive organs are located on the underside of the body. Males have two slits, one concealing the penis and one further behind for the anus. The female has one genital slit, housing the vagina and the anus. A mammary slit is positioned on either side of the female's genital slit. A recent study at the US National Marine Mammal Foundation revealed that dolphins are the only animals other than humans that develop a natural form of Type 2 Diabetes, which may lead to a better understanding of the disease and new treatments for both humans and dolphins.

Senses


Most dolphins have acute eyesight, both in and out of the water, and they can hear frequencies ten times or more above the upper limit of adult human hearing. Though they have a small ear opening on each side of their head, it is believed that hearing underwater is also if not exclusively done with the lower jaw, which conducts sound to the middle ear via a fat-filled cavity in the lower jaw bone. Hearing is also used for echolocation, which all dolphins have. It is believed that dolphin teeth function as an antenna to receive incoming sound and to pinpoint the exact location of an object. The dolphin's sense of touch is also well-developed, with free nerve endings densely packed in the skin, especially around the snout, pectoral fins and genital area. However, dolphins lack an olfactory nerve and lobes and thus are believed to have no sense of smell. They do have a sense of taste and show preferences for certain kinds of fish. Since dolphins spend most of their time below the surface, tasting the water could function like smelling, in that substances in the water can signal the presence of objects that are not in the dolphin’s mouth. Though most dolphins do not have hair, they do have hair follicles that may perform some sensory function. The small hairs on the rostrum of the Boto river dolphin are believed to function as a tactile sense possibly to compensate for the Boto's poor eyesight.



Behavior

See also: Whale surfacing behaviour Dolphins are often regarded as one of Earth's most intelligent animals, though it is hard to say just how intelligent. Comparing species' relative intelligence is complicated by differences in sensory apparatus, response modes, and nature of cognition. Furthermore, the difficulty and expense of experimental work with large aquatic animals has so far prevented some tests and limited sample size and rigor in others. Compared to many other species however, dolphin behavior has been studied extensively, both in captivity and in the wild. See cetacean intelligence for more details

Social behavior

Dolphins are social, living in pods of up to a dozen individuals. In places with a high abundance of food, pods can merge temporarily, forming a superpod; such groupings may exceed 1,000 dolphins. Individuals communicate using a variety of clicks, whistles and other vocalizations. They make ultrasonic sounds for echolocation. Membership in pods is not rigid; interchange is common. However, dolphins can establish strong social bonds. Dolphins will stay with injured or ill individuals, even helping them to breathe by bringing them to the surface if needed. This altruism does not appear to be limited to their own species however. The dolphin Moko in New Zealand has been observed guiding a female Pygmy Sperm Whale together with her calf out of shallow water where they had stranded several times. They have also been seen protecting swimmers from sharks by swimming circles around the swimmers or charging the sharks to make them go away. Dolphins also display culture, something long believed to be unique to humans (and possibly other primate species). In May 2005, a discovery in Australia found Indo-Pacific Bottlenose Dolphin (Tursiops aduncus) teaching their young to use tools. They cover their snouts with sponges to protect them while foraging. This knowledge is mostly transferred by mothers to daughters, unlike simian primates, where knowledge is generally passed on to both sexes. Using sponges as mouth protection is a learned behavior. Another learned behavior was discovered among river dolphins in Brazil, where some male dolphins use weeds and sticks as part of a sexual display. Dolphins engage in acts of aggression towards each other. The older a male dolphin is, the more likely his body is to be covered with bite scars. Male dolphins engage in such acts of aggression apparently for the same reasons as humans: disputes between companions and competition for females. Acts of aggression can become so intense that targeted dolphins sometimes go into exile as a result of losing a fight. Male Bottlenose Dolphins have been known to engage in infanticide. Dolphins have also been known to kill porpoises for reasons which are not fully understood, as porpoises generally do not share the same diet as dolphins and are therefore not competitors for food supplies.

Reproduction and sexuality


Dolphin copulation happens belly to belly and though many species engage in lengthy foreplay, the actual act is usually brief, but may be repeated several times within a short timespan. The gestation period varies per species; for the small Tucuxi dolphin, this period is around 11 to 12 months, while for the Orca the gestation period is around 17 months. They usually become sexually active at a young age, even before reaching sexual maturity. The age of sexual maturity varies by species and gender.

Dolphins are known to have sex for reasons other than reproduction, sometimes also engaging in homosexual behavior. Various species sometimes engage in sexual behavior including copulation with other dolphin species. Sexual encounters may be violent, with male dolphins sometimes showing aggressive behavior towards both females and other males. Occasionally, dolphins behave sexually towards other animals, including humans.

Feeding

Various methods of feeding exist among and within species, some apparently exclusive to a single population. Fish and squid are the main food, but the False Killer Whale and the Killer Whale also feed on other marine mammals.

One common feeding method is herding, where a pod squeezes a school of fish into a small volume, known as a bait ball. Individual members then take turns plowing through the ball, feeding on the stunned fish. Coralling is a method where dolphins chase fish into shallow water to more easily catch them. In South Carolina, the Atlantic Bottlenose Dolphin takes this further with strand feeding, driving prey onto mud banks for easy access. In some places, Orcas come to the beach to capture sea lions. Some species also whack fish with their fluke, stunning them and sometimes knocking them out of the water.

Reports of cooperative human-dolphin fishing date back to the ancient Roman author and natural philosopher Pliny the Elder. A modern human-dolphin partnership currently operates in Laguna, Santa Catarina, Brazil. Here, dolphins drive fish towards fishermen waiting along the shore and signal the men to cast their nets. The dolphins’ reward is the fish that escape the nets.

Vocalizations

Dolphins are capable of making a broad range of sounds using nasal airsacs located just below the blowhole. Roughly three categories of sounds can be identified: frequency modulated whistles, burst-pulsed sounds and clicks. Dolphins communicate with their whistles and burst-pulsed sounds, though the nature and extent of that ability is not known. At least some dolphin species can identify themselves using a signature whistle.The clicks are directional and are for echolocation, often occurring in a short series called a click train. The click rate increases when approaching an object of interest. Dolphin echolocation clicks are amongst the loudest sounds made by marine animals.

Jumping and playing

Dolphins occasionally leap above the water surface, sometimes performing acrobatic figures (e.g. the Spinner Dolphin). Scientists are not certain about the purpose(s) of the acrobatics. Possibilities include locating schools of fish by looking at above-water signs like feeding birds, communicating with other dolphins, dislodging parasites or simple amusement.

Play is an important part of dolphin culture. Dolphins play with seaweed and play-fight with other dolphins. At times they harass other local creatures, like seabirds and turtles. Dolphins enjoy riding waves and frequently surf coastal swells and the bow waves of boats, at times “leaping” between the dual bow waves of a moving catamaran. Occasionally, they playfully interact with swimmers.

Sleeping






Source:http://en.wikipedia.org/wiki/Dolphin